ABSTRACT

IN RECEN T YEAR S student s o f natura l an d o f artificia l population s hav e demon -strated enormou s geneti c heterogeneit y i n suc h population s an d hav e com e to realiz e tha t ther e ar e man y factor s tha t w i l l maintai n such geneti c hetero - geneity. Earlier , though th e classica l mathematician s o f th e subject , Fisher , Hal -

dane, an d Wright , ha d discusse d selectio n i n clines , density-dependen t factors ,

and th e effect s o f gene migratio n betwee n differen t adaptiv e peaks , mor e simple -

minded biologist s concluded fro m th e mathematic s tha t natura l population s migh t

be expecte d t o b e geneticall y rather uniform ; suc h variet y as the y envisage d wa s

regarded a s dependen t largel y on recurren t mutation .