ABSTRACT

Abstract .................................................................................................................... 67 Keywords .................................................................................................................68 4.1Introduction ....................................................................................................68 4.2Induction of Presoldier Differentiation by JH and JHA Application ............. 70

4.2.1Search for Effective JHs and JHAs .................................................... 70 4.2.2 Effects of JHA on Nasus and Frontal Gland Formation .................... 72

4.3 Organ Development during Presoldier Differentiation in N. takasagoensis ..... 74 4.3.1Formation of the Nasus Disk and Frontal Gland ................................ 74 4.3.2Regression of the Mandible ................................................................ 74

4.4Conclusion ...................................................................................................... 78 Acknowledgments .................................................................................................... 79 References ................................................................................................................ 79

arti„cial induction of soldier differentiation, it was shown that organ formation was involved in the organ-speci„c molecular mechanisms causing the epidermal folding (nasus disc), the epidermal invagination (frontal gland) and the epithelial programmed cell death (mandible). Moreover, the limb-patterning gene Distalless was involved in at least nasus formation. These results suggest that castespeci„c regulation of preexisting developmental mechanisms, triggered by JH, underlie morphological modi„cations during the caste differentiation of termites.