ABSTRACT

Acknowledgments....................................................................................................84 References................................................................................................................85

We view natural selection as producing strategies that approximately maximize an appropriate measure of fitness. Although the trajectories of the evolutionary process may be important, we concentrate on the endpoints of the process. We discuss a state-dependent approach to behavior in which an animal is characterized by state variables such as size, immunocompetence, and energy reserves. State-dependent reproductive value can be used to characterize evolutionary stability. This can be done even when there are intergenerational effects (phenotypic inheritance of traits) such as cultural transmission. We use an example of the coevolution of female mate choice rules and male trait value (the Fisher process) to show that it is possible to