ABSTRACT
Pronotum as wide as elytra basally, basal margin slightly to conspicuously bisinuate, unmargined; laterally narrowed toward apex; sides narrowly to broadly margined. Prosternum prominent, with intercoxal process extending laterally well behind coxae. Prothoracic coxal cavities closed internally and externally. Scutellum visible, rounded posteriorly. Mesosternum with intercoxal process short, broad; mesothoracic coxal cavities open laterally. Metasternum transverse. Legs moderately long; trochantins of prothoracic legs often partially exposed by slit-like extension of coxal cavity; prothoracic coxae globose, always well separated; mesothoracic coxae rounded, well separated, metathoracic coxae transverse, contiguous; trochanters small, triangular, heteromeroid; femora slender to slightly swollen; tibiae slender, sometimes spinose along outer edge; tibial spurs prominent; metathoracic tibiae sometimes modified in males. Tarsal formula 5-5-4 (rarely 4-4-4); tarsomeres 1-2 or 1-3 densely setose ventrally; penultimate tarsomere often highly reduced. Elytra slightly longer than wide, strongly convex, with fine to coarse rows of punctures; elytral intervals smooth and shining or punctulate; epipleural fold broad. Metathoracic wings usually fully developed with short radial cell; medial fleck present, undivided.
