ABSTRACT

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112NYMPHAEACEAE (incl. Cabombaceae, excl. Nelumbonaceae)

Genera

5–9 Cabomba, Euryale, Nuphar and Nymphaea (water lily), Victoria.

Species

50–90.

Distribution

Cosmopolitan.

Description of the family

Habit:

Perennial herbs, aquatic, found in fresh water. Rhizomes. Stem immersed.

Leaves:

Alternate, long-petiolate, simple, cordate or peltate, generally floating.

Inflorescence:

Flower solitary, large and showy.

Flower:

3–6 S / 3-n P / 3-n St / 2-n C. Long-pedicellate, large, spiralate, homoio- or heterochlamydeous, actinomorphic, polystemonous, hypogynous or epigynous, bisexual. Sepals often petaloid. Petals often indefinite resulting from the gradual transformation of stamens. Many lamellar stamens sometimes staminodal; anthers with longitudinal dehiscence. Ovary superior or inferior, multilocular, many free or connate carpels; stigma sessile, united and flat or in a disc above the carpels; placentation laminar, many ovules per locule, anatropous, bitegmic.

Fruit:

Fleshy capsule tough or woody. Small seeds often arillate; endosperm poorly developed, perisperm abundant.

Placement in the systems

• Engler:

Ranunculales

• Thorne:

Nymphaeanae-Nymphaeales

• Cronquist:

Magnoliidae-Nymphaeales

• Dahlgren:

Nymphaeiflorae-Nymphaeales

A, Floral diagram of the genus <italic>Nymphaea</italic> https://s3-euw1-ap-pe-df-pch-content-public-p.s3.eu-west-1.amazonaws.com/9780429187889/63dd0025-042c-4470-8ca7-4c71d437eeb2/content/fig101_112_1_B.tif"/> <target id="page_113" target-type="page">113</target> <italic>Nymphaea alba</italic>: B, habit; C, longitudinal section of the flower; D, detail of the flat stigma; E, cross-section of the ovary with parietal placentation, laminar; F, tip of an indefinite petal. https://s3-euw1-ap-pe-df-pch-content-public-p.s3.eu-west-1.amazonaws.com/9780429187889/63dd0025-042c-4470-8ca7-4c71d437eeb2/content/fig101_113_1_B.tif"/>

114ALISMATACEAE

Genera

12 Alisma (water plantain), Echinodorus, Sagittaria (arrowleaf).

Species

75.

Distribution

Cosmopolitan, well represented in the northern hemisphere. Found in aquatic and marshy environments.

Description of the family

Habit:

Aquatic herbs, rooted at the bottom of the bed. Rhizomes. Latiferous schizogenic ducts.

Leaves:

Basal, alternate, sagittate or linear to cordate; petiole well developed, sheathed.

Inflorescence:

Axillary.

Flower:

3 S / 3 P / (3-) 6 (-n) St / (3-) 6-n C. Heterochlamydeous, trimerous, actinomorphic, diplo- or polystemonous, hypogynous, bisexual. Petals caducous. Three to six stamens in a whorl or many stamens arranged in a spiral; anthers bilocular; pollen multiporous. Nectaries on the receptacle, at the base of floral parts or carpels. Ovary superior, carpels free; style terminal or gynobasic; stigma decurrent; placentation basal, ovule generally solitary, anatropous to amphitropous, bitegmic.

Fruit:

Achenes, rarely follicles dehiscent at the base. Seed non-endospermic.

Placement in the systems

• Engler:

Helobiae-Alismatineae

• Thorne:

Alismatanae-Alismatales

• Cronquist:

Alismatidae-Alismatales

• Dahlgren:

Alismatiflorae