ABSTRACT
Territoriality is one of the most important behavioural traits affecting the spatial organization of animal populations. Kruuk (1972) showed that spotted hyena (Crocuta crocuta) were territorial in one population but not in another. This difference was determined by the nature of their food supplies. Davies (1978) recognized territoriality where animals were spaced further apart than would be expected from a random occupation of suitable habitats. Territories are often considered as spatially stable and this is made explicit, for example, in Brown and Orians’ (1970) definition as ‘a fixed, exclusive area with the presence of defence that keeps out rivals’. Homerange is defined as ‘that area traversed by the individual in its normal activities of food gathering, mating and caring for young’ (Burt 1943). The concept has been refined by several authors (Mohr 1947; Jewell 1966; Baker 1978): home-range is not the whole area that an animal traverses during its lifetime (Jewell 1966), but rather the area over which an animal normally travels in pursuit of its routine activities.
