Lakes are extremely important storages of the earth’s surface freshwater, with the important ecosystem service functions that can continue the sustainable development of society and the economy. 1,2 However, eutrophication, acidification, heavy metal, oil, and pesticide pollution caused by human activities have continuously deteriorated the healthy status of lake ecosystems. Water quality in over half of the lakes around the world has been seriously polluted. If this trend continues, it not only affects human health and social-economic development, but also may cause the breakup of lake ecosystems. 3,4 Therefore, the studies on lake ecosystem health have important and practical significance for the restoration of lake ecosystem health and the maintenance of their ecological service functions.