ABSTRACT

Trees in a Pine Stand ................................................................................................ 233 6.5.3 Range Expansion of the Pinewood Nematode with Reference to the

Long-Distance Dispersal of M. alternatus ...................................................................... 234 6.5.4 Description of Regional Dispersal of M. alternatus and B. xylophilus Using

an Analytic Approach ...................................................................................................... 236 6.5.5 Description of Regional Dispersal of M. alternatus Using DNA Markers ..................... 236

The genus Monochamus Dejean belongs to the subfamily Lamiinae of the family Cerambycidae, which is one of the largest and most important among wood-boring families (Linsley 1961). Species from this genus are called “pine sawyers” because the larvae make a loud noise as they tunnel within the wood of host trees. They feed on conifers and broad-leaved trees. In Japan, some species feed on both (Kojima and Nakamura 2011). Many species attack dying, newly dead, or freshly cut trees of the genera Pinus L., Picea Mill., and Abies Mill., and are considered secondary pests.