Introduction Tourism is one of the world’s leading industries. In 2014, global tourism generated 9% of the world’s GDP, exports of US$ 1.4 trillion, provided one in eleven jobs (UNWTO, 2015), and generated approximately US$ 600 billion in direct in-country expenditures from protected areas visitation (Balmford et al., 2015). Through such activities, nature-related tourism has become an important component of biodiversity conservation (Morrison et al., 2012; Pegas et al., 2013; Steven, Castley & Buckley, 2013) and capacity building (Mbaiwa, 2011) efforts in developing nations.