ABSTRACT

Trees are long lived and must adapt to non-temporal ecological dynamics in their chosen ecosystem as well as those of the temporal plane. One implication is with evolutionary-derived ecosystem membership where membership is also a temporal affair. A positive effect may be temporally one-on-one or ecosystem related. From the successional phases, with variations, a modest theoretical base emerges which explain the dynamics of temporal silviculture. There are some general guidelines that relate intra-period complementarity to the inter-period version. If two species are complementary, growing well together, they will generally grow well as temporal neighbors although the temporal ordering may be important. The growth progression of a natural forest is one of overlapping successions of different species. In the temporal plane, light and water may be less of a factor in temporally separate monocultures or polycultures, bringing to the fore positive complementarity in mineral resources.