ABSTRACT
Pronotum convex with base wider than or subequal to elytral base and with or without tubercles, ridges, horns, or sulci. Elytra convex, with or without striae. Pygidium concealed by elytra. Scutellum exposed, triangular. Legs with coxae transverse, mesocoxae separated or contiguous; protibiae serrate on outer margin, apex with 1 spur; meso- and metatibia with ridges, apex with 2 spurs; tarsi 5-5-5; claws equal in size, simple; empodium present, extending beyond fifth tarsomere, with 2 setae.
