ABSTRACT
Pronotum large cowled or flat; shape subquadrate or anteriorly rounded; borders not margined (except Lyctinae and some
Xyloperthini, in which case margins either complete or basal, smooth or denticulate); cowled species with anterior dorsal surface rough, rasp-like, some with curved horns or hooks; pleural region broad; some flattened types (Lyctinae) with a median depression, or a median fovea; prosternum long to moderate in front of the coxae; intercoxal process wide to absent; procoxal cavities open or closed behind. Mesosternum moderate. Metasternum broad, long. Legs with the trochantins hidden; anterior coxae small, cylindrical to rounded, middle coxae round, nearly contiguous; hind coxae transverse, nearly contiguous to widely separate; trochanter obliquely or squarely attached to femur; femora broad to slender; tibiae slender, smooth, with single apical spur, tarsal formula 5-5-5, rarely 4-4-4 (Psoa), first tarsomere frequently very small, tarsomeres slender, simple. Scutellum small, triangular, oval, or quadrate. Elytra highly variable, usually coarsely punctate, with vague costae present, more rarely impunctate and plane; often with a distinct apical declivity variously modified with spines; epipleural fold obscure. Hind wing well developed, in fully-veined species with characteristic bostrichoid venation, a short r3 attached to the RP directly from the rp-mp2 cross-vein, a well-developed R cell; folding with normal transverse folds (Forbes 1926).
