ABSTRACT
Pronotum as wide or wider than head, broadly oval to subquadrate, borders mostly completely margined laterally, but margin may be present only at base or completely absent; surface smooth, punctate, or rugose; pleural region broad; prosternum very short, in some deeply excavated with mesosternum in front
for head; procoxal cavities externally open, internally closed. Mesosternum short; mesocoxae moderately separated to separated by more than one coxal width, laterally coxae closed by sternum. Metasternum broad, convex.
