ABSTRACT
Pronotum broad, evenly convex or depressed anteromedially, widest at or behind middle, lacking tubercles, horns or sulci, but some species with a low, transverse median ridge; mesothoracic spiracles primitive, slipper-shaped, with 2 intersegmentalia on each side. Scutellum exposed, subtriangular, narrowly to broadly rounded apically, lightly to densely pubescent. Elytra convex with variable sculpturing (costal striae lacking, feebly indicated, coriaceous, and/or strongly indicated); lateral margins rounded to sutural angle, sutural margins contiguous to apices. Pygidium exposed in both sexes. Legs with procoxae large, conical and prominent, procoxal cavities open; mesocoxae contiguous, prominent; protibiae strongly toothed on outer margin; meso- and metatibiae strongly ridged externally; tarsi simple and
subcylindrical, elongate, 5-5-5, tarsomeres 1-4 subequal in length, tarsomere 5 longer than preceding 2 together; empodium present, produced beyond apex of tarsomere 5, with 2-3 setae.
