Characteristics of TSV virions include a buoyant density of approximately 1.338 g mL−1 in CsCl, icosahedral morphology, a size of 31-32 nm in diameter, and a positive sense singlestranded RNA (ssRNA) genome that is polyadenylated at the 3′ end [16]. Therefore, the virus was initially classied as a member of the family Picornaviridae [5,16]. However, genomic characterization and sequence comparison revealed that TSV is closely related to the cricket paralysis-like virus [17]. Presently, TSV has been classied by the International Committee on Taxonomy of Viruses into the novel genus Aparavirus in a new family Dicistroviridae in the order Picornavirales [18].