Regarding a vegan culture, Darwinian descent with modification is covered in this chapter through dimensions like genetic, epigenetic, behavioral, and symbolic evolution. Standard texts with updated material are included. Selection on both the individual and group levels is addressed in terms of feeding and diet adaptations for survival and reproduction. Synergy, genetic control, and kin selection are covered in line with the evolution of behavior as they relate to veganism. Contrasts between wildlife environments and modern, industrial humans are made. Biogeography is examined in terms of veganism according to biota, energy budgets, and allometry with consideration given to the differences between great apes and humans. Evolutionary psychology is discussed in terms of vegan behavior, especially disgust responses, along with the evolution of diet. Brain size is debated in terms of evolutionary diets and other selection pressures.